The Mission Dispatcher is a structure in the DLCsGenesis: Part 1 and Genesis: Part 2. They are used to start Missions within the Genesis Simulations, and are also the primary means of transferring tames and items across servers.
Each Dispatcher has an icon above it representing the type of Mission offered when interacted with. However, the Mission "One Tough Moeder" stands out, as it appears larger with an exclamation mark icon, both above and at the screen.
In Part 2
Any dispatcher found on the map can be used to start any Mission.
The Admin Blink Rifle can destroy Mission Dispatchers, so be careful when shooting near them as they do not respawn.
You can spawn generic (without mission) Mission Dispatcher for Genesis: Part 1 with console command: cheat summon missiondispatcher_multiusepylon_c
You can spawn Mission Dispatcher for Genesis: Part 2 with console command: cheat summon missiondispatcher_gen2_c